Ferns, lycophytes, and their extinct free-sporing relatives
Dryopteris carthusiana
Dryopteris carthusiana(Vill.) H. P. Fuchs
Family: Dryopteridaceae
[Aspidium callipteris Wilms, moreAspidium spinulosum (O.F.Muell.) Sw., Aspidium spinulosum f. anadenium B.L.Rob., Aspidium spinulosum var. elevatum A. K. H. Braun ex Döll, Dryopteris austriaca subsp. spinulosa O.F.Müller ex H. Schinz & Thellung, Dryopteris austriaca var. spinulosa (O.F.Müll.) Fiori, Dryopteris eu-spinulosa (Aschers.) Fomin, Dryopteris spinulosa var. elevata (A. K. H. Braun ex Döll) Vollmann, Dryopteris spinulosa var. exaltata (Lasch) Vollmann, Dryopteris spinulosa var. glandulosa Gilbert, Dryopteris spinulosa var. patuloides Christ, Filix spinulosa (O.F.Muell.) Farw., Filix-mas spinulosa , Lastrea dilatata var. linearis C. C. Babington, Lastrea spinosa (Schrank) Newm., Lastrea spinulosa var. elevata (A. K. H. Braun ex Döll) J. T. I. B. Syme & N. E. D. Br., Lophodium spinosum (Schrank) Newm., Nephrodium jordanii Rouy, Nephrodium spinulosum (O.F.Muell.) Strempel, Nephrodium spinulosum f. anadenium (B.L.Rob.) Clute, Nephrodium spinulosum var. elevatum (A. K. H. Braun ex Döll) Murr, Nephrodium spinulosum var. swarzii F. W. Schultz, Polypodium carthusianum Vill., Polypodium filix-femina var. spinosa F. G. Weis, Polypodium spinulosum (Hillebr.) Bak., Polypodium tanacetifolium Hoffm., Polystichum angustatum Schum., Polystichum callipteris (Ehrh.) Bernh., Polystichum conifolium Schum., Polystichum pseudocristatum Schur, Polystichum spinosum (F. G. Weis) Roth, Polystichum spinulosum (O.F.Muell.) Lam. & DC., Polystichum spinulosum subsp. spinulosum , Thelypteris spinulosa (O.F.Muell.) Nieuwl.]
Rhizome horizontal, short-creeping; lvs deciduous, essentially glabrous except for the chaffy, concolorous pale brown scales that beset at least the lower part of the petiole and sometimes also part of the rachis; petiole mostly a quarter to a third as long as the blade; blade 2-5+ dm, a third to half as wide, broadest a little below the middle, bipinnate- pinnatifid to tripinnate, with mostly 10-15 pairs of pinnae below the pinnatifid tip; pinnae (at least the lower ones) mostly oblique, the basal pinnule of the lower side of the lowest pinna longer than the one next to it and to twice (thrice) as long as the subopposite upper pinnule; ultimate segments finely spinulose- toothed; sori midway between the midvein and the margins; 2n=164. Moist or wet woods and swamps; interruptedly circumboreal, in Amer. s. to S.C., Ark., and Wash. (D. spinulosa; D. austriaca var. s.)
Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.