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Notholaena standleyi (Kümmerle) Maxon  
Family: Pteridaceae
[Cheilanthes hookeri (Kümmerle) Domin, moreCheilanthes standleyi (Kümmerle) Mickel, Chrysochosma hookeri (Domin) Kümmerle, Chrysochosma standleyi Kümmerle, Cincinalis hookeri (Lowe) J. Sm., Hemionitis hookeri (E.J.Lowe) Christenh., Notholaena hookeri E.J. Lowe]
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Patrick Alexander
  • FNA
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Michael D. Windham in Flora of North America (vol. 2)
Stem scales strongly bicolored, margins brown, broad and well defined, thin, entire. Leaves 5--30 cm. Petiole brown, equal to or somewhat longer than blade, rounded adaxially, glabrous except for a few scales at base. Blade broadly pentagonal, deeply pinnatifid but not fully pinnate above base (distal pinnae connected by narrow wing of green tissue), 1--2 times longer than wide, abaxially with conspicuous yellowish farina, scales absent, adaxially glabrous; basal pinnae much larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged. Ultimate segments sessile, broadly adnate to costae; segment margins slightly recurved, rarely concealing sporangia. Sporangia containing 32 or 16 spores. 2 n = 60. Sporulating late spring--fall. Rocky slopes and cliffs, on a variety of substrates including granite and limestone; 300--2100 m; Ariz., Colo., N.Mex., Okla., Tex.; Mexico. The low numbers of spores per sporangium in Notholaena standleyi are apparently not associated with apogamy because all plants thus far analyzed are sexual diploids (M. D. Windham, unpublished data). D. S. Siegler and E. Wollenweber (1983) identified three chemotypes in this species correlated with substrate specificity and subtle variations in farina color. These three 'races' occupy different portions of the geographic range, and further investigation may indicate that they deserve formal taxonomic recognition. A report of Notholaena standleyi from Nevada (D. B. Lellinger 1985) must be considered suspect because it is disjunct, and the closest populations in Arizona represent a different chemotype.

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Leslie Landrum
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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Patrick Alexander
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The PCC, and this data portal, were made possible by funding from the National Science Foundation’s Advancing Digitization of Biological Collections (ADBC) program, grant numbers 1802504, 1802352, 1802134, 1802033, 1802270, 1802255, 1802239, 1802446, 1802305.
The pteridoportal taxonomic thesaurus is based on the Checklist of Ferns and Lycophytes of the World, generously provided by Michael Hassler.